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They prey on such small aquatic animals as insects and worms. Like other members of the family, darters possess two dorsal fins.
Their size ranges from 2. Some darters are among the most brightly coloured fishes in North America, the males becoming especially colourful during the spring breeding season.
Spawning habits vary. Some species scatter or bury their eggs and abandon them; in other species, the males establish nests and guard the eggs until hatched.
Numerous darter species are becoming rare, and several are listed as threatened or endangered in the Red Data Book. With an increasing number of nomads settling down in recent decades, this cultural heritage is in danger of being lost.
On the other hand, as evidenced by the etymology of "anhinga" detailed above, the Tupi seem to have considered the anhinga a kind of bird of ill omen.
The genus Anhinga was introduced by the French zoologist Mathurin Jacques Brisson in with the anhinga or American darter Anhinga anhinga as the type species.
However, in its first documented use as an English term in , it referred to an Old World darter. Ever since, it has also been used for the modern genus Anhinga as a whole.
This family is very closely related to the other families in the suborder Sulae , i. Cormorants and darters are extremely similar as regards their body and leg skeletons and may be sister taxa.
In fact, several darter fossils were initially believed to be cormorants or shags see below. Some earlier authors included the darters in the Phalacrocoracidae as subfamily Anhingina', but this is nowadays generally considered overlumping.
However, as this agrees quite well with the fossil evidence  some unite the Anhingidae and Phalacrocoracidae in a superfamily Phalacrocoracoidea.
The Sulae are also united by their characteristic display behavior, which agrees with the phylogeny as laid out by anatomical and DNA sequence data.
While the darters' lack of many display behaviors is shared with gannets and that of a few with cormorants , these are all symplesiomorphies that are absent in frigatebirds , tropicbirds and pelicans also.
Like cormorants but unlike other birds, darters use their hyoid bone to stretch the gular sac in display.
Whether the pointing display of mates is another synapomorphy of darters and cormorants that was dropped again in some of the latter, or whether it evolved independently in darters and those cormorants that do it, is not clear.
The male raised-wing display seems to be a synapomorphy of the Sulae; like almost all cormorants and shags but unlike almost all gannets and boobies, darters keep their wrists bent as they lift the wings in display, but their alternating wing-waving, which they also show before take-off, is unique.
That they often balance with their outstretched wings during walking is probably an autapomorphy of darters, necessitated by their being plumper than the other Sulae.
The Sulae were traditionally included in the Pelecaniformes , then a paraphyletic group of " higher waterbirds ". The supposed traits uniting them, like all-webbed toes and a bare gular sac , are now known to be convergent , and pelicans are apparently closer relatives of storks than of the Sulae.
Hence, the Sulae and the frigatebirds — and some prehistoric relatives — are increasingly separated as the Suliformes , which is sometimes dubbed "Phalacrocoraciformes".
There are four living species of darters recognized, all in the genus Anhinga ,  although the Old World ones were often lumped together as subspecies of A.
They may form a superspecies with regard to the more distinct anhinga: . Extinct "darters" from Mauritius and Australia known only from bones were described as Anhinga nana "Mauritian darter" and Anhinga parva.
In the former case, however, the remains are larger than those of the geographically closest extant population of long-tailed cormorants on Madagascar : they thus might belong to an extinct subspecies Mauritian cormorant , which would have to be called Microcarbo africanus nanus or Phalacrocorax a.
The Late Pleistocene Anhinga laticeps is not specifically distinct from the Australasian darter; it might have been a large paleosubspecies of the last ice age.
The fossil record of the Anhingidae is rather dense, but very apomorphic already and appears to be lacking its base. The other families placed in the Phalacrocoraciformes sequentially appear throughout the Eocene , the most distinct — frigatebirds — being known since almost 50 Ma million years ago and probably of Paleocene origin.
With fossil gannets being known since the mid-Eocene c. Fossil Anhingidae are known since the Early Miocene ; a number of prehistoric darters similar to those still alive have been described, as well as some more distinct genera now extinct.
The diversity was highest in South America , and thus it is likely that the family originated there. Some of the genera which ultimately became extinct were very large, and a tendency to become flightless has been noted in prehistoric darters.
Their distinctness has been doubted, but this was due to the supposed "Anhinga" fraileyi being rather similar to Macranhinga , rather than due to them resembling the living species: .
Prehistoric members of Anhinga were presumably distributed in similar climates as today, ranging into Europe in the hotter and wetter Miocene. With their considerable stamina and continent-wide distribution abilities as evidenced by the anhinga and the Old World superspecies , the smaller lineage has survived for over 20 Ma.
As evidenced by the fossil species' biogeography centered around the equator , with the younger species ranging eastwards out of the Americas, the Hadley cell seems to have been the main driver of the genus' success and survival: .
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Archived from the original on 10 July Retrieved 18 July Archived from the original on 2 March Archived from the original on 22 February Flight International.
Archived from the original on 31 December Retrieved 31 December Archived from the original on 11 February Retrieved 11 February Archived from the original on 23 February Retrieved 25 September In Spitzer, Cary ed.
The Avionics Handbook. Archived from the original on 29 July Denel Dynamics.